Redescription of the Cranial Morphology of Mariliasuchus Amarali, and Its Phylogenetic Affinities (crocodyliformes, Notosuchia)

The cranial morphology of Mariliasuchus amarali, a poorly known notosuchian from the Late Cretaceous of Southeastern Brazil, is redescribed based on new material. Its phylogenetic affinities within Crocodylomorpha are evaluated through a parsimony analysis involving 46 taxa and 198 characters. Mariliasuchus is nested well inside the clade Notosuchia, as the sister group of Comahuesuchus, a derived notosuchian from the Late Cretaceous of Argentina. Both taxa share the following unambiguous synapomorphies: ventral half of the lacrimal tapering posteroventrally, not contacting or only slightly contacting the jugal; presence of a large foramen on the lateral surface of the anterior part of the jugal; presence of procumbent premaxillary and anterior dentary alveoli; and ectopterygoids that do not participate of the palatine bar. The presence of procumbent premaxillary teeth, specialized tooth crown morphology, and fore–aft jaw movements suggests that this group presented complex jaw movements related to specialized feeding habits.