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The Wooley Creek batholith is a Late Jurassic, arc-related, calc-alkaline plutonic complex in the Klamath Mountain province of California. Post-emplacement tilting and erosion have exposed ~12 km of structural relief. The complex consists of an older (~159.1 Ma) lower zone (pyroxenite to tonalite) assembled by piecemeal emplacement of many magma batches, a younger (~158.2 Ma) upper zone (quartz diorite to granite), and a transitional central zone. In the lower zone, pyroxenes are too Fe rich to be in equilibrium with a melt whose composition was that of the host rock. Mass-balance calculations and simulations using rhyolite-MELTS indicate that these rocks are cumulates of pyroxenes and plagioclase ± olivine and accessory apatite and oxides. Percentages of interstitial melt varied from ~7.5–83%. The plagioclase/pyroxene ratios of cumulates vary considerably among the most mafic rocks, but are relatively uniform among quartz diorite to tonalite. This near-constant ratio results in compositional trends that mimic a liquid line of descent. In the upper zone, bulk-rock compositional trends are consistent with differentiation of andesitic parental magmas. Upward gradation from quartz dioritic to granitic compositions, modeled via mass-balance calculations and rhyolite-MELTS simulations, indicate that structurally lower parts of the upper zone are cumulates of hornblende and plagioclase ± biotite and accessory minerals, with 37–80% trapped melt. In contrast, the structurally higher part of the upper zone represents differentiated magma that escaped the subjacent cumulates, representing differentiated melt fractions remaining from 92–54%. The ratio of cumulate plagioclase/(plagioclase + mafic minerals) is ~0.48 among upper-zone cumulates, mimicking a liquid line of descent.