Oribatid mite fossils from a terrestrial Devonian deposit near Gilboa, New York

Fourteen oribatid mite fossils have been recovered from a terrestrial Devonian deposit near Gilboa, New York; they allow an approximate doubling of the known age of the group. Two species are represented by specimens complete enough to allow descriptions. Protochthonius gilboa n. gen. and sp. is the earliest derivative known member of the Enarthronota, and comprises the monobasic new family Protochthoniidae. Devonacarus sellnicki n. gen. and sp. also represents a monobasic new family, Devonacaridae; it may also be an early derivative enarthronote mite, but its relationships are uncertain. Two other species are represented at the site, but meaningful descriptions are not possible with available material.