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Feeding environment and feed accessibility influence the dairy cow's response to the ration and forage composition. Fiber content, physical form, and fermentability influence feeding behavior, feed intake, and overall cow metabolic and lactational responses to forage. It is possible to vary eating time of lactating dairy cattle by over 1 h/d by changing dietary silage fiber content, digestibility, and particle size. Optimizing silage particle size is important because excessively long particles increase the necessary chewing to swallow a bolus of feed, thereby increasing eating time. Under competitive feeding situations, excessively coarse or lower fiber digestibility silages may limit DMI of lactating dairy cows due to eating time requirements that exceed available time at the feed bunk. Additionally, greater silage particle size, especially the particles retained on the 19-mm sieve using the Penn State Particle Separator, are most likely to be sorted. Silage starch content and fermentability may influence ruminal propionate production and thereby exert substantial control over meal patterns and feed consumption. Compared with silage fiber characteristics, relatively little research has assessed how silage starch content and fermentability interact with the feeding environment to influence dairy cow feeding behavior. Finally, voluminous literature exists on the potential effects that silage fermentation end products have on feeding behavior and feed intake. However, the specific mechanisms of how these end products influence behavior and intake are poorly understood in some cases. The compounds shown to have the greatest effect on feeding behavior are lactate, acetate, propionate, butyrate, ammonia-N, and amines. Any limitation in the feeding environment will likely accentuate the negative response to poor silage fermentation. In the future, to optimize feeding behavior and dry matter intake of silage-based diets fed to dairy cattle, we will need to consider the chemical and physical properties of silage, end products of silage fermentation, and the social and physical components of the feeding environment.