Taxonomic and nomenclatorial notes on the Catoptria coulonellus (Duponchel) species group with description of five new species from Western Balkans (Lepidoptera: Pyraloidea, Crambidae)

The extensive study of the original descriptions and of rich material allowed us to review the Catoptria coulonellus species group, with its species C. coulonellus (Duponchel, 1836) sp. reinst., C. combinella ([Denis & Schiffermüller], 1775) = C. petrificella (Hübner, 1796) syn. nov., C. hannemanni Alberti, 1967, C. orientellus (Herrich-Schäffer, 1850), C. spodiellus (Rebel, 1916), C. trichostomus (Christoph, 1858), C. lythargyrella (Hübner, 1796), C. domaviellus (Rebel, 1904), C. nikai sp. nov., C. richteri sp. nov., C. kasyi Błeszyński, 1960, C. korab sp. nov., C. plat sp. nov., C. sarplanina sp. nov., and C. biformellus (Rebel, 1893). Neotypes of C. combinella, C. petrificella, C. orientellus and C. lythargyrella are designated to fix the identity of these names. All species are fully illustrated, females of C. hannemanni for the first time. Finally, DNA barcodes are provided for a selection of available species and applied for an integrative species delimitation.