The systematics and nomenclature of Tethysuchia (Archosauria: Crocodylomorpha) under the International Code of Phylogenetic Nomenclature

Tethysuchian evolutionary relationships have been intensively investigated, in particular those of Dyrosauridae. In order to provide the most up to date phylogenetic hypothesis, we performed a phylogenetic analysis including Brachiosuchus kababishensis and Wadisuchus kassabi, with goniopholidids and tethysuchians (= pholidosaurids and dyrosaurids). The dataset we propose here is based on several previous analyses (e.g. Jouve et al. 2005a, b, 2008; Hastings et al. 2015; Young et al. 2017) and includes 225 characters and 44 taxa. Compared with previous iterations of this character list (Jouve 2020; Jouve and Jalil 2020; Jouve et al. 2021), several definitions have been clarified, and the final character in the matrix is new. Two new taxa have been added, Brachiosuchus kababishensis and Wadisuchus kassabi, and numerous characters and other taxa have been rescored.