Aphanogmus fumipennis Thomson 1858

Aphanogmus fumipennis species group Diagnosis. Randomly sized areolae on head and mesosoma absent. Median mesoscutal sulcus present, transscutal articulation present. Lateral carina on mesoscutellum absent, posterior lamella of mesoscutellum absent. Lateroventral mesopleural carina present. Posterior mesosomal comb present, scattered setae on posteriomedian metacoxa absent, setal patch on proximal metacoxa absent, posteriodistal spine on metacoxa present (Fig. 2). Transverse carina of syntergite present, submedial setiferous patches on synsternite absent. Stigmal vein longer than pterostigma marginal length. Male genitalia. Dorsal apodeme of penisvalva present. Ventromedian apodeme of aedeagus absent. Medioventral area of gvc not folded: Parossicular seta oriented ventrally, visible in lateral view. Aedeagus not widened. Harpe usually finger-shaped, often with basomedian and median spine (e. g., Figs 4 D, 27 D) and sometimes with lamella-like extension along median margin (e. g., Figs 15 D, 25 D). Female terminalia. Tergo-valviferal articulation (tva) on the posterior margin of the first valvifer (1 vf) located in middle, not angled. First valvula (1 vv) gradually tapering distally. Second valvula (2 vv) slightly shorter than first valvula. Basal line of the second valvifer (bl) and anterior section of dorsal flange of the second valvifer (asf) distinct. Terebra (trb) slightly longer than ovipositor assembly. Ovipositor length virtually equal to mesonotum length in lateral view (Fig. 3). Species included. Aphanogmus bispinatus Salden, Mikó & Peters, sp. nov., A. crispy Salden, Mikó & Peters, sp. nov., A. digitangulus Salden, Staverløkk, Mikó & Peters, sp. nov., A. fumipennis Thomson, 1858, sp. inq., A. fraterculus Salden & Peters, 2023, A. hafti Salden, Mikó & Peters, sp. nov., A. hamatogonus Salden, Staverløkk, Mikó & Peters, sp. nov., A. harpagodalis Salden, Mikó & Peters, sp. nov., A. karlazani Salden, Mikó & Peters, sp. nov., A. kintrishi Salden, Mikó & Peters, sp. nov., A. latiharpus Salden & Peters, sp. nov., A. pilosicoxa Salden & Peters, 2023, A. simbai Salden & Peters, 2023, A. tkheliharpus Salden, Mikó & Peters, sp. nov., A. trigoharpus Salden, Mikó & Peters, sp. nov., A. vashlovani Salden, Mikó & Peters, sp. nov., A. ventricoharpus Salden, Staverløkk, Mikó & Peters, sp. nov., A. zefranki Salden, Mikó & Peters, sp. nov. Distribution. Afrotropics: Kenya. Palaearctic: Czech Republic, Georgia, Germany, Hungary, Norway, Slovakia, Sweden, and Switzerland. Biology. Unknown. Remarks. The posterior mesosomal comb is the main diagnostic character and allows quick identification of the A. fumipennis species group. It consists of two rows of densely packed setae interlocking at their distal parts and bordering a smooth, concave area along the posterior metacoxa. The posterodistal spine of the metacoxa marks the terminal point of the comb. On the posterolateral surface of the metanoto-propodeo-mesopecto-metapectal complex, densely packed setae continue (Fig. 2). Aphanogmus was previously divided into three species groups by Evans et al. (2005), based on the presence or absence of two morphological characters: the median mesoscutal sulcus (“ mesoscutal median furrow ” in Evans et al. (2005)) and the transverse carina of the syntergite (“ gastral basal carina ”). In the clavicornis group, both characters are absent; in the tenuicornis group the median mesoscutal sulcus is absent while the transverse carina of the syntergite is present; and in the fumipennis group both characters are present. These species groups have been adopted in recent taxonomic work (Matsuo et al. 2016; Salden and Peters 2023; Mollaei et al. 2025). The present A. fumipennis species group diagnosis significantly expands and refines the previous diagnosis of the fumipennis group by Evans et al. (2005).