Variable Density Planting: Using Marigolds as a Model System to Describe a Silvicultural Approach to Increase Structural Diversity

There is limited information on the effects of irregular-spaced and clumped planting on forest production and structural diversity. We explored Tagetes patula L. development as a model system to demonstrate stand development under varying planting patterns, and conceptualized in the context of Pseudotsuga menziesii (Mirb.) Franco mesic production forests of the Pacific Northwestern USA. Two variable planting patterns, clumps of 24 plants and of four plants, were compared to square regular patterns in replicate growing boxes. Spatial patterns were compared post-planting and at maturation, along with stand-level metrics and final dry weights, and stand structural statistics were used to compare production and diversity. The clumped spatial structure of 24-plant clumps was maintained to maturity. Groups of four plants maintained clumping at small scales and regular patterns at larger scales. Initial Regular-Square spacing remained at 2 cm at maturity but became indistinguishable from random patterns at larger scales. There was (1) overall greater mean directional index for the Large-Clumped patterns and greater spatial complexity indices for both clumped patterns, (2) greater social class (size) mean mingling index for small clumps, and (3) higher mean dominance index and mean differentiation index and lower crown volume complexity and height-to-diameter ratios for Regular-Square spacing. The structural complexity was accompanied by limited differences in dried weights by plant tissue (total weight, stem, leaf, flower weight) or plant biometric parameters (stem straightness, crown ratio, crown volume, number of leaves and flowers). The results from irregular planted marigold stand development are discussed in the context of increasing forest stand complexity, potentially without compromising productivity.