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Odorrana yangi sp. nov. Figs 2, 3, 4 Type material. Holotype. • KIZ 2025143, adult male, collected by Chao Bu on 5 April 2025 from Muba Village, Tianpeng Town, Funing County, Wenshan Prefecture, Yunnan Province, China (23°22'8"N, 105°34'3"E, 710 m a. s. l.). Paratypes. • KIZ 2025123, adult female, • KIZ 2025124 and KIZ 2025144 – KIZ 2025148, six adult males, all the same collection data as that of the holotype. Etymology. The specific epithet is in commemoration of the renowned Chinese herpetological taxonomist Prof. Dr Datong Yang, who recently passed away. We suggest “ Yang’s Odorous Frog ” as the common English name and “ 杨氏臭蛙 ” as the common Chinese name. Diagnosis. Body size small in males (SVL 43.5–46.9 mm), females approximately two times size of males; head length greater than head width; nostril closer to tip of snout than to eye; tympanum relatively large in males (TD / ED 0.53–0.63); heels overlap when hindlimbs flexed at right angles to axis of body, tibiotarsal articulation reaching tip of snout when hindlimb stretched forward; relative lengths of fingers III> IV> I> II; dorsal surface relatively smooth, skin on dorsum shagreened, with some indistinct flat tubercles; dorsolateral fold absent; anterior dorsum green with evenly distributed, small, irregular-shaped, black blotches, posterior dorsum greyish brown with evenly distributed, large, irregular-shaped, black blotches; ventral surface white without distinct spots or patches; external vocal sacs and nuptial pads present in adult males. Description of holotype. Adult male, body size small (SVL 46.4 mm); head length greater than head width (HL / HW 1.12); snout obtuse, pointed in dorsal view and rounded in profile, projecting beyond lower jaw; nostril closer to tip of snout than to eye (NED / NSD 1.15); canthus rostralis distinct, blunt; loreal region concave; internarial distance greater than interorbital distance (IND / IOD 1.49); snout length greater than eye diameter (SL / ED 1.17); pineal body distinct; supratympanic fold distinct; eye moderate (ED / HDL 0.38); tympanum relatively large (TYD / HDL 0.20, TYD / ED 0.53), round; vomerine teeth distinct, two oblique ridges between two choanae; choanae small, rounded; tongue large, cordiform, posterior notched; vocal sac openings small, on floor of mouth in each corner; external vocal sacs below angle of mouth on each side. Forelimb robust; fingers long, relative length III> IV> I> II; tips of fingers I and II slightly expanded, tips of fingers III and IV obviously expanded into disks; lateral fringes and webbing on fingers absent; subarticular tubercles small and distinct, oval, formula 1, 1, 2, 2; supernumerary tubercles absent; a large oval thenar tubercle on ventral finger I; metacarpal tubercles indistinct; nuptial pad on dorsal finger I. Hindlimb long (HLL / SVL 1.75), tibia longer than thigh (TL / THL 1.06); toes moderately long, relative length IV> V> III> II> I; all toe tips expanded into disks; fringes on out edges of toes I and VI present; toe webbing formula I 1–11 / 3 II 1 – 1 ½ III 1 – 2 IV 2 – 1 V; subarticular tubercles small and distinct, rounded, formula 1, 1, 2, 3, 2; supernumerary tubercles absent; inner metatarsal tubercle small, oval; outer metatarsal and tarsal fold absent. Dorsal skin shagreened with some indistinct flat tubercles, skin on flanks slightly granular, dorsolateral fold absent, ventral skin smooth. Colouration of holotype in life. Dorsal head and anterior dorsum green with evenly distributed, small, irregular-shaped, black blotches, gradually transitioning to yellowish brown with evenly distributed, large irregular-shaped, black blotches posteriorly; dorsal limbs yellowish brown, with brownish-black transverse stripes; upper lib yellow with brownish-black vertical stripes; tympanum flesh-coloured, translucent, with a small, opaque, light-yellow dot at centre; iris light copper-coloured, with black mesh-like pattern; flank yellow, with some large, irregular-shaped, brownish-black spots; ventral surface of head and body white; vocal sac flesh-coloured; ventral surface of limbs light purple. Variations. Morphological measurements of all type specimens are given in Table 2 and the photographs of two paratypes in life are shown in Fig. 4. The male paratypes are similar to the holotype in morphology and coloration except for the pineal body is almost invisible in some individuals. The female paratype is similar to the males in coloration but significantly larger than the males in body size. In addition, the forelimbs are relatively slender, and the nuptial pad and vocal sac are absent in the female paratype. Natural history. The specimens of the new species were all collected on the rocks or tree branches along a large river at night. Banks of the river are karst landforms. No eggs or tadpoles were found during the survey. Odorrana tiannanensis (Yang & Li, 1980) was found to be distributed sympatric with the new species. Distribution. The new species is currently known only from its type locality in Funing County, Wenshan Prefecture, Yunnan Province, China (Fig. 5). Comparisons. Odorrana yangi sp. nov. differs from O. absita (Stuart & Chan-ard, 2005), O. amamiensis (Matsui, 1994), O. aureola Stuart, Chuaynkern, Chan-ard & Inger, 2006, O. banaorum (Bain, Lathrop, Murphy, Orlov & Ho, 2003), O. bolavensis (Stuart & Bain, 2005), O. chloronota (Günther, 1876), O. confusa, O. damingshanensis, O. exiliversabilis Li, Ye & Fei, 2001, O. gigatympana (Orlov, Ananjeva & Ho, 2006), O. graminea (Boulenger, 1900), O. hosii (Boulenger, 1891), O. heatwolei (Stuart & Bain, 2005), O. indeprensa (Bain & Stuart, 2006), O. khalam (Stuart, Orlov & Chan-ard, 2005), O. leporipes (Werner, 1930), O. livida (Blyth, 1856), O. macrotympana (Yang, 2008), O. monjerai (Matsui & Jaafar, 2006), O. montivaga (Smith, 1921), O. morafkai (Bain, Lathrop, Murphy, Orlov & Ho, 2003), O. narina (Stejneger, 1901), O. nasica (Boulenger, 1903), O. nasuta Li, Ye & Fei, 2001, O. orba (Stuart & Bain, 2005), O. supranarina (Matsui, 1994), O. tiannanensis, O. tormota (Wu, 1977), O. utsunomiyaorum (Matsui, 1994), O. versabilis, and O. yentuensis Tran, Orlov & Nguyen, 2008 by dorsolateral folds being absent (vs present). Odorrana yangi sp. nov. differs from O. andersonii, O. bacboensis, O. cangyuanensis (Yang, 2008), O. chapaensis (Bourret, 1937), O. geminata Bain, Stuart, Nguyen, Che & Rao, 2009, O. grahami (Boulenger, 1917), O. ishikawae (Stejneger, 1901), O. jingdongensis Fei, Ye & Li, 2001, O. junlianensis Huang, Fei & Ye, 2001, O. kuangwuensis (Liu & Hu, 1966), O. lungshengensis (Liu & Hu, 1962), O. margaretae (Liu, 1950), O. nanjiangensis, O. mawphlangensis (Pillai & Chanda, 1977), O. mutschmanni Pham, Nguyen, Le, Bonkowski & Ziegler, 2016, O. swinhoana (Boulenger, 1903), O. tianmuii, and O. wuchuanensis (Xu, 1983) by having a smaller body size in males (SVL <47 mm vs SVL> 48 mm). Odorrana yangi sp. nov. differs from O. calciphila, O. concelata, O. feii, O. leishanensis Li, Chen, Su, Liu, Tang & Wang, 2024, O. liboensis, and O. lipuensis by vocal sacs being present in males (vs absent). Odorrana yangi sp. nov. differs from O. anlungensis (Liu & Hu, 1973), O. hejiangensis, O. huanggangensis, O. ichangensis, O. kweichowensis, and O. schmackeri by tibiotarsal articulation reaching snout tip (vs reaching nostril or between eye and nostril); from O. dulongensis by head length being greater than head width (vs head length being smaller than head width); from O. fengkaiensis and O. sangzhiensis by tibiotarsal articulation reaching snout tip (vs beyond snout tip); from O. hainanensis by having a pair of external vocal sacs in males (vs a pair of internal vocal sacs); from O. splendida Kuramoto, Satou, Oumi, Kurabayashi, & Sumida, 2011 by tibiotarsal articulation reaching snout tip (vs reaching anterior corner of eye); and from O. sudianensis and O. yizhangensis Fei, Ye & Jiang, 2007 by having different relative length of fingers (III> IV> I> II vs III> I> IV> II in O. sudianensis and III> IV> II> I in O. yizhangensis).